Solved What does the comma in a package filename mean?

Having carefully read the Porter's Handbook chapter on specifying version numbers, I remain puzzled by the name /var/cache/pkg/m4-1.4.17_1,1.txz. What does the ,1 mean?

At first I thought it was a sort of European-style decimal point in the PORTEPOCH. But that can't be it, because I also notice /var/cache/pkg/bison-2.7.1,1.txz, which has no PORTEPOCH but still has that ,1.

What's up with that? What does it mean?
 
Well, that’s PORTREVISION, isn’t it? I’m not sure…

EDIT: I think that this syntax is used when PORTREVISION and PORTEPOCH must be shown together.
 
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